#NEXUS [written Thu Feb 06 10:50:20 EST 2025 by Mesquite version 3.61 (build 927) at HUIT-FAS-OEB-MMapalo/140.247.97.54] BEGIN TAXA; TITLE Taxa; DIMENSIONS NTAX=71; TAXLABELS Tubiluchus_troglodytes_Priapulida Priapulus_caudatus_Priapulida Ottoia_prolifica_Archeopriapulida Cricocosmia_jinningensis Aysheaia_pendunculata Onychodictyon_ferox Onychodictyon_gracilis Diania_cactiformis Xenusion_auerswaldae Paucipodia_inermis Microdictyon_sinicum Cardiodictyon_catenulum Hallucigenia_sparsa Hallucigenia_fortis Hallucigenia_hongmeia Orstenotubulus_evamuellerae Luolishania_longicruris Collinsium_ciliosum Facivermis_yunnanicus Ovatiovermis_cribratus Collinsovermis_monstrousus Collins_monster_Emu_Bay Acinocricus_stichus Antennacanthopodia_gracilis Euperipatoides_Peripatopsidae Metaperipatus_blainvillei_Peripatopsidae Epiperipatus_biolleyi_Peripatidae Florarctus_bellahelenae_Halechiniscidae Batillipes_pennaki_Batillipididae Actinarctus_doryphorus_Tanarctidae Coronoarctus_sonne_Coronarctidae Anisonyches_diakidius_Anisonychidae Neoarctus_primigenius_Neoarctidae Stygarctus_ayatori_Stygarctidae Styraconyx_takeshii_Styraconyxidae Archechiniscus_bahamensis_Arcechniscidae Neostygarctus_oceanopolis_Neostygarctidae Renaudarctus_fossorius_Renaudarctidae Echiniscus_testudo_Echiniscidae Proechiniscus_hanneae_Echiniscidae Bryodelphax_mareki_Echiniscidae Echiniscoides_sigismundi_Echiniscoididae Isoechiniscoides_sifae_Echiniscoididae Carphania_fluviatilis_Carphaniidae Oreella_mollis_Oreellidae Milnesium_eurystomum_Milnesiidae Isohypsibius_prosostomus_Isohypsibiidae Hypsibius_dujardini_Hypsibiidae Macrobiotus_hufelandi_Macrobiotidae Eohypsibius_nadjae_Eohypsibiidae Megadictyon_haikouensis Siberion_leanicus Jianshanopodia_decora Hadranax_augustus Kerygmachela_kierkegaardi Pambdelurion_whittingtoni Opabinia_regalis Anomalocaris_canadensis Peytoia_nathorsti Hurdia_victoria Aegirocassis_benmoulae Lyrarapax_unguispinus Schinderhannes_bartlesi Fuxianhuia_xiaoshibaensis Chengjiangocaris_kunmingensis Leanchoilia_superlata Alalcomenaeus_sp. Misszhouia_longicaudata Kuamaia_lata Limulus_polyphemus Triops_cancriformis ; END; BEGIN CHARACTERS; TITLE Character_Matrix; DIMENSIONS NCHAR=94; FORMAT DATATYPE = STANDARD RESPECTCASE GAP = - MISSING = ? SYMBOLS = " 0 1 2 3 4 5"; CHARSTATELABELS 1 Paired_appendages, 2 Anterior_region_covered_by_sclerites, 3 Head_shield_formed_by_fused_cephalic_segments_, 4 'Isolated dorsal sclerite associated with eye-stalks', 5 Shape_of_dorsal_isolated_sclerite, 6 Extent_of_coverage_of_dorsal_isolated_sclerite_on_head_, 7 'Isolated lateral sclerites, forming tri-partite carapace', 8 'Nature of post-ocular (post-protocerebral) body appendages ', 9 Protocerebral_limb_pair_with_arthrodial_membranes_, 10 Protocerebral_appendage_pair_fused_, 11 Nature_of_protocerebral_appendage_fusion, 12 'Presence of spines along the protocerebral appendage (i.e., not the terminal part)', 13 'Number of spine/spinule series on protocerebral frontal appendage', 14 'Coplanar spine/spinule series in protocerebral frontal appendages', 15 Presence_of_multifurcate_terminal_spines_in_protocerebral_appendage, 16 'Nature of post-ocular lobopodous inner branch', 17 Mouth_opening_orientatio, 18 One_or_more_pairs_of_appendages_located_anteriorly_relative_to_the_mouth_opening, 19 Radially_symmetrical_circumoral_mouth_structures, 20 Nature_of_radial_circumoral_structures, 21 Structure_of_mouth_apparatus, 22 Papillate_circumoral_structure, 23 Eyes, 24 Eye_attachment, 25 Types_of_eyes, 26 Epidermal_segmentation, 27 Dorsal_integument_sclerotized_and_connected_by_arthrodial_membranes, 28 Sternites_connected_by_arthrodial_membranes, 29 'Neck-like constriction on lobopodous trunk', 30 Annulations, 31 Organization_of_trunk_annulation, 32 Presence_of_epidermal_evaginations, 33 Proportions_of_epidermal_trunk_evagination, 34 Trunk_epidermal_evaginations_with_acute_distal_termination, 35 Acute_distal_termination_in_epidermal_evagination_is_curved, 36 Sclerotization_of_epidermal_evaginations, 37 Sclerites_consist_of_a_stack_of_constituent_elements, 38 Maximum_number_of_primary_dorsal_epidermal_specializations_above_each_leg_pair, 39 'Dorsal sclerotized spine-like evaginations of variable length along the body', 40 Spacing_between_dorsolateral_epidermal_specializations_along_longitudinal_body_axis, 41 Papillae_on_trunk_annulations, 42 'Serially repeated mid-gut glands', 43 Trunk_exites, 44 Exite_organization, 45 'Exites/lanceolate dorsal blades associated with dorsolateral flaps', 46 'Exite/setal blade distribution', 47 'Dorsal flaps/exites fused with endopod into biramous appendage', 48 'Antero-posteriorly compressed protopodite with gnathobasic endites in post-deutocerebral appendage pair', 49 Secondary_structures_on_the_first_ambulatory_lobopodous_limbs, 50 Organization_of_secondary_structures_on_first_ambulatory_lobopodous_limbs, 51 Secondary_structures_on_the_second_ambulatory_lobopodous_limbs, 52 Organization_of_secondary_structures_on_second_ambulatory_lobopodous_limbs, 53 'Finger-like elements in distal tip of limbs', 54 'Secondary structures on posterior-most ambulatory lobopodous limbs ', 55 'Organization of secondary structures on posterior-most ambulatory lobopodous limbs', 56 Terminal_claws_on_trunk_limbs, 57 Organization_of_terminal_claws, 58 Number_of_claws_on_trunk_limbs, 59 Symmetry_of_fused_claws, 60 Differentiated_distal_foot_in_lobopodous_trunk_limbs, 61 Hypertrophied_set_of_anterior_body_flaps, 62 Strengthening_rays_in_lateral_flaps, 63 Posterior_tapering_of_lateral_flaps, 64 Anterior_sets_of_reduced_lateral_flaps, 65 'Lobopodous limbs differentiated into two batches of multiple anterior/long and posterior/short limbs', 66 Number_of_anterior_morphologically_differentiated_elongated_limbs, 67 Appendages_comprise_15_or_more_podomeres, 68 Limbless_posterior_extension_of_the_lobopodous_trunk_beyond_last_appendage_pair, 69 Posterior_appendage_as_appendicular_tail, 70 Claws_on_posterior_appendages_directed_anteriad, 71 Dorsal_heart, 72 Dorsal_condensed_brain, 73 Number_of_neuromeres_integrated_into_the_dorsal_condensed_brain, 74 Mouth_innervation_relative_to_brain_neuromeres, 75 General_organization_of_ventral_nerve_cord, 76 Ventral_nerve_cord_with_morphologically_discrete_condensed_hemiganglia_connected_by_median_commissures, 77 Paired_nerve_cord_with_median_interpedal_commissures, 78 Nerve_cord_with_orthogonal_organization, 79 'Orthogonal nerve cord with complete ring-commissures', 80 Segmental_leg_nerves_shifted_anteriorly_relative_to_appendages_following_parasegmental_organization, 81 Paired_segmental_leg_nerves_, 82 Regularly_spaced_peripheral_nerves_running_entire_length_of_nerve_cord, 83 Lateral_cirri, 84 Median_cirrus, 85 Telescopic_legs, 86 Placoids, 87 Eversible_mouthparts, 88 Number_of_trunk_limbs, 89 Lunules_at_the_base_of_fused_claws, 90 Bulbous_pharynx, 91 Caudal_appendage, 92 Caudal_hooks, 93 Body_size, 94 Internalized_lobopodous_limbs ; MATRIX Tubiluchus_troglodytes_Priapulida 00--------------0-10-00--00-0100--------10----------------------------00--00-11--100-01--1100- Priapulus_caudatus_Priapulida 00--------------0-10-00--00-0100--------10----------------------------00--00-11--100-01--1101- Ottoia_prolifica_Archeopriapulida 00--------------0-10-00--00-0100--------10----------------------------??????-??--?00-01--1011- Cricocosmia_jinningensis 00--------------0-10-00--00-010100-101-010----------------------------?????????--?00-?1--1011- Aysheaia_pendunculata 10-----000-10-10000--10--00-0100--------100-----1010010104-0----0--001????????????000?11-1--10 Onychodictyon_ferox 10-----000-11100000--010000-0111010101-0100-----1111011111-0----0--001????????????000??1-1--10 Onychodictyon_gracilis 10-----000-11?00100--00--00-0111010101-0100-----????0??111-0----0--100????????????000??1-?--10 Diania_cactiformis 10-----000-0--00000--00--00-011100-1?0-0100-----1111011????0----0--10?????????????000?01-?--10 Xenusion_auerswaldae 10-----000-????0?00--00--00-011100-??1-01?0-----????????????----0--10?????????????000??1-?--10 Paucipodia_inermis 10-----000-0--00000--00--00-0100--------000-----0-0-00-111-0----0--100????????????000?01-0--10 Microdictyon_sinicum 10-----000-0--00100--00--00-011100-101-0000-----0-0-00-111-0----0--100????????????000?01-0--10 Cardiodictyon_catenulum 10-----000-0--00?0????10?00-011100-1?1-0000-----0-0-00-111-0----0--100????????????000?01-1--10 Hallucigenia_sparsa 10-----000-0--00100--0???00-00-111011100-00-----0-0-00-111-0----0--00?????????????000?01-0--10 Hallucigenia_fortis 10-----000-0--00100--010?00-01111111?100000-----0-0-00-111-0----0--100????????????000?01-?--10 Hallucigenia_hongmeia 1??????0???????0?????????00-?1111111?110000-----????0??110-0----??-10???????????????0?01-?--1? Orstenotubulus_evamuellerae 1??????0???????0?????????00-?1111????1????0-----????????????----??-?????????????????????-?--?? Luolishania_longicruris 1100---000-0--00100--010?00-01111111?211100-----1111011110-0----10-100????????????000?01-0--10 Collinsium_ciliosum 1100---000-0--00100--00--00-011111111411100-----111100-110-0----11-100????????????000?01-?--10 Facivermis_yunnanicus 10-----000-????0000--010000-0110--------100-----11110110---0----10-10-????????????000?01-0--10 Ovatiovermis_cribratus 10-----0???????000????10000-0110--------000-----111100-110-0----11-001????????????000?01-?--10 Collinsovermis_monstrousus 1100---000-0--00000--00--00-01111111?211000-----111100-110-0----11-000????????????000?01-?--10 Collins_monster_Emu_Bay 110????0???????0?????????00-01111111?21???0-----11110??110-0----10-???????????????000??1-?--10 Acinocricus_stichus 1??????0???????0?????????00-01111111?511?00-----11110???????----10-???????????????0?0??1-?--10 Antennacanthopodia_gracilis 10-----000-0--00?0????10000-01?0--------0?0-----1111?11????0----0--10?????????????000?01-1--10 Euperipatoides_Peripatopsidae 10-----000-0--00110--010000-0100--------100-----1111011111-1----0--100111310111011000001-1--10 Metaperipatus_blainvillei_Peripatopsidae 10-----000-0--00110--010000-0100--------100-----1111011111-1----0--100111310111011000001-1--10 Epiperipatus_biolleyi_Peripatidae 10-----000-0--00110--010000-0100--------100-----1111011111-1----0--100111310111011000001-1--10 Florarctus_bellahelenae_Halechiniscidae 10-----000-0--00100--00--00-00-0---------00-----1010110113-0----0--0010100111??110111110-1--01 Batillipes_pennaki_Batillipididae 10-----000-0--00100--010000-00-0---------00-----10101100---0----0--001-100111??110111110-1--01 Actinarctus_doryphorus_Tanarctidae 10-----000-0--00100--010000-00-0---------00-----1010110113-0----0--0010100111??110111110-1--01 Coronoarctus_sonne_Coronarctidae 10-----000-0--00000--00--00-00-0---------00-----1010010103-0----0--0010100111??110110110-1--01 Anisonyches_diakidius_Anisonychidae 10-----000-0--00000--010000-00-0---------00-----0-0-01010(2 3)-0----0--0010100111??1101101?0-1--01 Neoarctus_primigenius_Neoarctidae 1100---000-0--00100--00--00-00-100-10?-0-00-----1010010102-0----0--0010100111??1101111?0-1--01 Stygarctus_ayatori_Stygarctidae 1100---000-0--00100--00--00-00-100-10?-0-00-----0-0-010103-0----0--0010100111??110111110-1--01 Styraconyx_takeshii_Styraconyxidae 10-----000-0--00100--00--00-00-0---------00-----1010110113-0----0--0010100111??1101111?0-1--01 Archechiniscus_bahamensis_Arcechniscidae 10-----000-0--00000--010000-00-0---------00-----100-(0 1)101(0 1)3-0----0--0010100111??110110100-1--01 Neostygarctus_oceanopolis_Neostygarctidae 1100---000-0--00100--00--00-00-100-10?-0-00-----0-0-111113-0----0--0010100111??110111110-1--01 Renaudarctus_fossorius_Renaudarctidae 1100---000-0--00100--00--00-00-100-10?-0-00-----0-0-110113-0----0--0010100111??1101111?0-1--01 Echiniscus_testudo_Echiniscidae 1100---000-0--00000--010000-00-100-10?-0-00-----100-010103-0----0--001010011110110100110-1--01 Proechiniscus_hanneae_Echiniscidae 1100---000-0--00000--010000-00-100-10?-0-00-----100-010103-0----0--0010100111??110100110-1--01 Bryodelphax_mareki_Echiniscidae 1100---000-0--00000--010000-00-100-10?-0-00-----100-010103-0----0--0010100111??110100110-1--01 Echiniscoides_sigismundi_Echiniscoididae 10-----000-0--00000--010000-00-0---------00-----0-0-010104-0----0--0010100111??110100110-1--01 Isoechiniscoides_sifae_Echiniscoididae 10-----000-0--00000--010000-00-0---------00-----1010010104-0----0--0010100111??110100110-1--01 Carphania_fluviatilis_Carphaniidae 10-----000-0--00000--00--00-00-0---------00-----101000-103-0----0--0010100111??110101110-1--01 Oreella_mollis_Oreellidae 10-----000-0--00000--00--00-00-0---------00-----100-010103-0----0--0010100111??110101110-1--01 Milnesium_eurystomum_Milnesiidae 10-----000-0--000011-110000-00-0---------00-----0-0-00-1(0 1)3-0----0--0010100111??110000010-1--01 Isohypsibius_prosostomus_Isohypsibiidae 10-----000-0--000013-0???00-00-0---------00-----0-0-00-12300----0--0010100111??1100001?001--01 Hypsibius_dujardini_Hypsibiidae 10-----000-0--00000--010000-00-0---------00-----0-0-00-12300----0--0010100111101100001?001--01 Macrobiotus_hufelandi_Macrobiotidae 10-----000-0--000011-010000-00-0---------00-----0-0-00-12310----0--001010?111??11000011011--01 Eohypsibius_nadjae_Eohypsibiidae 10-----000-0--000011-00--00-00-0---------00-----0-0-00-12300----0--001010?111??11000011011--01 Megadictyon_haikouensis 10-----000-10-10001??00--00-0100--------010-----1111011????0----0--???????????????000?01-1--10 Siberion_leanicus 10-----000-1???010???10--00-0100--------1?0-----???????????0----0--10?????????????000?01-?--10 Jianshanopodia_decora 10-----00??10-?0001??00--00-0100--------010-----1111011????0----0--11?????????????000?01-1--10 Hadranax_augustus 1??????0???????0?????????00-?11100-0-3-01?0-----0-0-?0-????0----0--?????????????????0??1-?--10 Kerygmachela_kierkegaardi 10-----000-10-100011-010100-011100-0-3-01110100-0-0-00-0---000000--10-?100????????000?01-1--10 Pambdelurion_whittingtoni 10-----000-10-?01012000--00-01???????????110100-0-0-00-0---000000--??-????????????00??01-1--10 Opabinia_regalis 10-----001010-1021??-011110-00-0---------110110-0-0-00-0---000000--11-????????????000?01-0--10 Anomalocaris_canadensis 110100001101101111121011100-10-0---------110010-0-0-00-0----01110--11-????????????000?01-?--10 Peytoia_nathorsti 1101100011010-1111121011100-00-0---------110110-0-0-00-0----01110--00-????????????000?01-?--10 Hurdia_victoria 110110101??10-11111210111?0-00-0---------?10110-0-0-00-0----01010--11-????????????000?01-?--10 Aegirocassis_benmoulae 110110101??10-11?1???????00-00-0---------?10110-0-0-00-0----011?0--00-??????????????0?01-?--10 Lyrarapax_unguispinus 1101000011010-1111121011100-10-0---------010010-0-0-00-0----1?110--11-?1001???????000?01-1--10 Schinderhannes_bartlesi 1??????0110110??111210111???0??????????????????-????????????1???0--11-??????????????0?01-?--10 Fuxianhuia_xiaoshibaensis 110101-10110--0-210--0111110-0-0---------011--10------------------111-1121????????000?01-0--1- Chengjiangocaris_kunmingensis 110101-10110--0-210--0111110-0-0---------011--10------------------111-????11?????1000?01-0--1- Leanchoilia_superlata 1110---10110--0-210--011111?-0-0---------111--11------------------010-????????????000?01-0--1- Alalcomenaeus_sp. 1110---10110--0-210--011111?-0-0---------111--11------------------010-?1211100-??0000?01-?--1- Misszhouia_longicaudata 111101-10110--0-210--0111111-0-0---------112--11------------------010-????????????000?01-0--1- Kuamaia_lata 111101-10110--0-210--0111111-0-0---------112--11------------------010-????????????000?01-?--1- Limulus_polyphemus 1110---10110--0-210--0101111-0-0---------114--11------------------010-11211100-100000001-0--1- Triops_cancriformis 1110---10110--0-210--0101111-0-0---------113--11------------------010-11221100-100000001-0--1- ; END; BEGIN ASSUMPTIONS; TYPESET * UNTITLED = unord: 1- 94; END; BEGIN MESQUITECHARMODELS; ProbModelSet * UNTITLED = 'Mk1 (est.)': 1- 94; END; Begin MESQUITE; MESQUITESCRIPTVERSION 2; TITLE AUTO; tell ProjectCoordinator; timeSaved 1738857020998; getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa; tell It; setID 0 2261567199411382633; tell It; setDefaultOrder 0 69 70 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 65 66 67 33 34 35 36 37 38 39 40 41 42 43 44 45 68 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64; attachments ; endTell; endTell; getEmployee #mesquite.charMatrices.ManageCharacters.ManageCharacters; tell It; setID 0 5886607512410878984; tell It; setDefaultOrder 0 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 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setFontIncAnnot 0; setFontIncExp 0; setSize 1289 988; setLocation 11 23; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.ibeam; endTell; setActive; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.ibeam; colorCells #mesquite.charMatrices.NoColor.NoColor; colorRowNames #mesquite.charMatrices.TaxonGroupColor.TaxonGroupColor; colorColumnNames #mesquite.charMatrices.CharGroupColor.CharGroupColor; colorText #mesquite.charMatrices.NoColor.NoColor; setBackground White; toggleShowNames on; toggleShowTaxonNames on; toggleTight off; toggleThinRows off; toggleShowChanges on; toggleSeparateLines off; toggleShowStates on; toggleReduceCellBorders off; toggleAutoWCharNames on; toggleAutoTaxonNames off; toggleShowDefaultCharNames off; toggleConstrainCW on; toggleBirdsEye off; toggleShowPaleGrid off; toggleShowPaleCellColors off; toggleShowPaleExcluded off; togglePaleInapplicable on; togglePaleMissing off; toggleShowBoldCellText off; toggleAllowAutosize on; toggleColorsPanel off; toggleDiagonal on; setDiagonalHeight 80; toggleLinkedScrolling on; toggleScrollLinkedTables off; endTell; showWindow; getWindow; tell It; forceAutosize; endTell; getEmployee #mesquite.charMatrices.AlterData.AlterData; tell It; toggleBySubmenus off; endTell; getEmployee #mesquite.charMatrices.ColorByState.ColorByState; tell It; setStateLimit 9; toggleUniformMaximum on; endTell; getEmployee #mesquite.charMatrices.ColorCells.ColorCells; tell It; setColor Red; removeColor off; endTell; getEmployee #mesquite.categ.StateNamesStrip.StateNamesStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.AnnotPanel.AnnotPanel; tell It; togglePanel off; endTell; getEmployee #mesquite.charMatrices.CharReferenceStrip.CharReferenceStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.QuickKeySelector.QuickKeySelector; tell It; autotabOff; endTell; getEmployee #mesquite.charMatrices.SelSummaryStrip.SelSummaryStrip; tell It; showStrip off; endTell; getEmployee #mesquite.categ.SmallStateNamesEditor.SmallStateNamesEditor; tell It; panelOpen true; endTell; endTell; endTell; endTell; end;